Studies of plant-pathogen interactions at the molecular level
Stess-induced DNA changes in plants
A survey and study of the plant pathogens reducing yields of indigenous food crops, and the transmission and control of these pathogens
Reproductive biology of seed plants
The isolation and identification of bioactive compounds from medicinal plants
Seed science
Development of Arbuscular mycorrhizal fungus inoculum and its practical use
Studies on the ephemerals of Namakwaland
Phytosociology, vegetation dynamics and conservation of the southern Kalahari
Phytosociology of the grassland and savanna biomes of South Africa
Systematics, biogeography and utilisation of the Southern African flora
Production of disease tolerant forest trees
Development of an oyster mushroom production industry in South Africa
Plant population and vegetation dynamics